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Germination
Early growth and tillering
Stem elongation
Wheat apex development
Flowering and fertilisation
Wheat ear pictures
Anthesis: floret pictures
Micrographs of fertilisation
Pollen release
Grain growth 1 to 4 days
Grain growth 4 to 10 days
Cell layers inside a grain
Grain filling 11 to 16 days
Grain filling 17 to 21 days
Grain filling 21 to 30 days
Development of the embryo
Dry down 30 to 40 days
How we grew the plants
Photography and microscopy
DAYS = days after flowering
![]() | The antipodal cells are fully formed, within the embryo sac, before the double fertilization. These cells are very large and are thought to provide the protein synthesizing machinery to support the 'free nuclear division' phase of the endosperm. The embryo sac is enclosed in a complex of laminated cell layers (some layers have parted in this section). The Cell Layers inside the Grain are described in detail in a separate section. |
![]() | High power longitudinal section of the embryo sac. The diploid zygote and the triploid nucleus formed at the second polar event are still both close to the micropyle. The antipodal cells are higher up in the embryo sac on the ventral side. |
![]() | Within a few hours of the union of male and female gametes the triploid fusion nucleus has divided, but remains near the zygote. |
![]() | One day after the double fertilization event. The zygote remains close to the micropyle, it may not have divided yet. The first endosperm nuclei, produced from the triploid fusion nucleus, have migrated toward the centre of the embryo sac. There have been several rounds of nuclear division without the formation of cell walls; this is the phase of 'free nuclear division'. |
![]() | The grain in longitudinal section at fertilization. The solid part of the carpel, the maternal pericarp, surrounds the embryo sac. The products of the double fertilization event are still close together. The antipodal cells are seen near the ventral wall. |